The brother of Sean Collier, the campus police officer killed three days after the attack on the Boston Marathon, remembered him Tuesday as having a strong moral compass — down to a belief that “you can’t kill a bug.”

Andrew Collier testified on the second day of the penalty phase of the trial of Dzhokhar Tsarnaev, who prosecutors say should be put to death for planning and executing the marathon bombings with his brother.

“It’s still a huge loss,” Andrew Collier said. “It is something that will affect my family for the rest of our lives.”

Sean Collier, an MIT police officer, was killed April 18, 2013, during the manhunt for the marathon suspects. His boss, MIT Police Chief John DiFava, remembered Collier as one of the best officers he has seen in decades of law enforcement.

“He always did the right thing,” DiFava testified. “Peer pressure did not work with Sean.”

On Monday, a federal prosecutor said that Tsarnaev was “determined to be America’s worst nightmare” and was unrepentant.

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